BIOL 1082 Lecture Notes - Lecture 12: Transpiration, Symplast, Plasmodesma

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19 Sep 2017
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The cell walls of guard cells are uneven in thickness, and the cellulose microfibrils are oriented in a direction that causes the guard cells to bow outward when turgid. This bowing outward increases the size of the pore b/w the guard cells. When the cells lose water and become flaccid, they become less bowed, and the pore closes. Stomata open when guard cells actively accumulate k+ from neighboring epidermal cells. The flow of k+ across the plasma membrane of the guard cell is coupled to the generation of a membrane potential by proton pumps. Stomatal opening correlates w/ active transport of h+ out of the guard cell. The resulting voltage drive k+ into the cell through specific membrane channels. The absorption of k+ causes the water potential to become more turgid as water enters through osmosis. Stomatal closing results from a loss of k+ from guard cells to neighboring cells, which leads to an osmotic loss of water.

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