BIOL 201 Lecture Notes - Absorption Spectroscopy, Carbon Fixation, Light-Independent Reactions

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19 Apr 2012
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L4. Photosynthesis I
Photosynthetic organisms capture light
energy (the ultimate source of biological energy) and
use it to:
Transfer H atoms from water to
acceptor molecules
Form molecular oxygen
Synthesize ATP from ADP and
phosphate
Transfer H atoms from acceptors to
carbons derived from CO2 to form glucose
(process known as carbon fixation)
Net reaction:
6 H2O + 6 CO2 C6H12O6 + 6 O2
Reduction of CO2
Oxidation of H2O
Endergonic reaction driven by
absorption of light
Light reactions:
2 H2O + 2 NADP+ + 3 ADP + 3 Pi
O2 + 2 H+ + 2 NADPH + 3 ATP
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In thylakoid
Calvin cycle (carbon fixation):
CO2 + 2 H+ + 2 NADPH + 3 ATP
CH2O + H2O + 2 NADP+ + 3ADP + 3 Pi
In stroma
Chloroplast structure
Outer membrane
Permeable to small molecules
Inner membrane
Transporters for phosphate and
sucrose precursors
Stroma (Calvin cycle/carbon fixation)
Enzymes that catalyze CO2 fixation
and starch synthesis
Thylakoid (light reactions)
Absorption of light by chlorophyll
Synthesis of ATP, NADPH
Electron transport
Stomata
Allow gas exchange: CO2 in, O2 out
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