ZOO 4070 Lecture Notes - Lecture 5: Inclusive Fitness, Parasitism

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Any behaviour that results in a direct itness cost to the emiter and a gain in the recipient. If behaviour evolves through natural selecion, how can altruisic behaviour spread: hamilton (1936-2000) It is generally accepted that the behaviour characterisic of a species is just as much a product of evoluion as the morphology . Problem: altruism is not easily explained by natural selecion. G > codes for altruism g > codes for selishness under standard geneic models, g cannot evolve. For an altruisic gene to evolve in a populaion: b > c, r x b > c rb) c >0 o a. 1: signiicance put behaviour into mathemaical framework created explicit predicions: can be applied not only to family but between and sets of individuals or groups redeines the concept of itness and introduces kin selecion theory. Assumpions: no guarantee of the number of ofspring, the quality of the males are the same.

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