Digestive Tract.docx

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Ecology & Evolutionary Biology
De Iuliis

Lecture 15 Digestive Tract JAN. 26  Digestive tract - tube delivering + absorbing food - things that are not absorbed will be removed from body  Visceral musculature = Muscle surrounding the gut that does the motion of transporting. This muscle produces wave-like contractions from mouth to anus, pushing food down the tube (=Peristalsis)  Reverse peristalsis = causes vomiting  No gravity is involved in digestion.  GI tract = produces mechanical digestion + chemical digestion where small bits of food are further broken down at the molecular level to be absorbed through wall of gut.  Absorption = getting the stuff across through wall of gut + enter the circulatory cardiovascular stream. Until this happens, the food is not part of us. The inside of GI tract is just a tube and it goes right through us. So inside of it is not actually technically part of our body. For that reason you have to get absorption. Need to have nutrients.  Gut can be divided into various parts. - Higher vertebrates – has segmented along the gut, uniform across vertebrates - Lower vertebrates – merging of pattern 1) Human: foregut + midgut + hindgut. This system doesn’t work well for majority of vertebrates 2) Shark: No distinction b/w small + large intestines, there is a single valvular intestine. 3) In some fish (e.g. lungfish): there is no stomach 4) In cyclostomes: Everything behind post-pharyngeal region is a simple tube.  There is a great deal of variation. The point is that condition is not comparable to humans or mammals in general. So there happens to be a landmark that is almost always present in vertebrates w/ very few exceptions, that is the constriction at the beginning of intestine called pylorus.  Pylorus = use that as a landmark to subdivide digestive tract into foregut & hindgut. That is a much easier way to look at the system in further developments.  In foregut we have: pharynx, esophagus, stomach - Pharynx: area of throat where gill slits are in the shark as well as in embryos - Distinction b/w esophagus & stomach is gross morphology: Consider esophagus to be a fairly thin pipe leading to a stomach (sac like structure). But the boundary b/w 2 is not always distinct. (e.g. in shark, you cannot see distinction until you cut + know difference internally. Stomach has rugae (longitudinal ridges) + esophagus (papillae). Therefore has a clear distinction internally but not externally.  Hindgut: follows pylorus that has valve called pyloric sphincter which can be opened & closed & therefore can control the passage of material from stomach (from foregut) to the hindgut. In hindgut we can usually but not always recognize various specializations. 1) Small intestine: duodenum, jejunum, ileum (no sharp distinction either morphologically or internally, it’s a tube of similar diameter throughout)  Duodenum: first loop of small intestine + loops towards the left & has part of pancreas adhering to surface + makes another turn & call it jejunum. 2) Large intestine 3) Rectum - In b/w small & large intestines or in vicinity b/w them, we often get out-pocketing that is a blind-ended pouch = Cecum - Cecum (plural ceca) – a small out-pocketing at the proximal end of the colon (e.g. b/w small & large intestine)  Functions of foregut: - Esophagus: we defined it as a thin pipe but sometimes it is not, and actually a tube where food is being ingested to where food will be treated. In vertebrates, there is ancestrally primitively very little digestion associated with the whole foregut but particularly with esophagus. - Stomach: develops as a large sac to store food. Being a part of foregut there is very little chemical digestion. In mammalian condition, we have a sharp distinction b/w esophagus and stomach. Esophagus being thin pipe and stomach as sac-like structure. In fish, not distinct. Fish have gill + pharyngeal region those are important part of foregut. Pharynx is very anterior end of GI tract (therefore part of foregut) but once we lose, pharynx becomes narrower & esophagus becomes a well developed pipe. Birds have a crop (a sac like ventral extension of esophagus, therefore in throat area; used as food storage).  Stomach becomes important when you need a place to dump food into. In cyclostomes having ancestral or primitive filter-feeding mechanism, the foods drivels back into GI tract at a slow rate. Let the food stay in stomach until it can be fed to intestine at a reasonable rate. That is what pyloric sphincter does controlling passage of material in stomach (storage area) & feed intestine at a reasonable rate. - We have almost no chemical digestion
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