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University of Toronto St. George
Human Biology

NROB60 Study Package Week 3 Homira OsmanWeek 3 Lecture May 20 2008Lecture TopicsLecture IIIReadings Development of the Nervous System Pages 187 to 192 Differentiation of the Midbrain Differentiation of the Hindbrain Differentiation of the Spinal Cord Putting the Pieces Together Classification Pages 48 to 49 The Prototypical Neuron Pages 26 to 46Lecture Summaries L3Differentiation of the Mesencephalic Vesicle MidbrainVery little development compared to ForebrainDorsal surface becomes the Tectum roofFloor becomes the Tegmentum CSFfilled space in between constricts into the Cerebral Aqueduct o The aqueduct connects rostrally with the third ventricle of the diencephalon o Because it is small and circular in cross section the cerebral aqueduct is a good landmark for identifying the midbrainDifferentiation of the MidbrainTectum Differentiates into superior colliculus and inferior colliculus Superior colliculus receives direct input from the eye o Controls eye movement o Cranial nerves III and IV NROB60 Study Package Week 3 Homira Osman Inferior colliculus receives direct input from the earRelay station en route to the thalamusTegmentum Most colourful regionsContains both the substantia nigra black substance and the red nucleus o Both are involved in control of voluntary movementOther cell groups have axons that project widely throughout CNS o Regulate consciousness mood pleasure and painStructureFunction Relationships of the MidbrainDiverseConduit for information passing from the spinal cord to the forebrain and vice versaAxons descending from cerebral cortex to brain stem and spinal cord ex corticospinal tractDamage results in loss of movement to opposite sideContribute to o Sensory systemso Control of movement o Several other functionsDifferentiation of the Rostral Hindbrain Hindbrain differentiates into three structureso Cerebellum Develops from rostral part of hindbrain o Pons Develops from rostral part of hindbrain o Medulla oblongata Develops from caudal part of hindbrainCalled Medulla CSFfilled tube becomes the fourth ventricleContinuous with cerebral aqueduct of the midbrain The rostral hindbrain differentiates into the cerebellum and ponsThe cerebellum is formed by the growth and fusion of the rhombic lipsThe CSFfilled space at the core of the hindbrain is the fourth ventricle
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