BIOL 005B Lecture Notes - Lecture 17: Gastrovascular Cavity, Hemolymph, Cellular Respiration

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11 May 2018
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BIOL 005B Lecture 17: Circulation
All organisms must exchange materials with the environment. The exchanges of gases
used in cellular respiration (O2 and CO2) is one of the most important processes in an
animal’s physiology
Most gases move easily through membranes by diffusion
Although diffusion requires no energy, it is a random process and is relatively
slow
Therefore, diffusion is fine for single celled and very small animals
Larger animals must build distributive systems for the transport of gas and other
materials
Distributive systems take advantage of convection (bulk flow
) through
circulating blood to carry materials from one place to another in the body
quickly and efficiently
Circulatory systems link exchange surfaces with cells throughout the body
In small and/or thin animals, cells can exchange materials directly with the
surrounding medium
In most animals, transport systems connect the organs of exchange with the body
cells
Most complex animals have internal transport systems that circulate fluid
Diffusion is only viable if it is fast enough
to support cellular and organismal
processes
In almost all animals that distance is less than 10 nm and effective over only one
or two cell layers
Gastrovascular Cavities
Simple animals, such as cnidarians, have a body wall that is only two cells
thick. It encloses a gastrovascular cavity - here diffusion might be possible
This cavity functions in both digestion and distribution of substances
throughout the body
Cnidarians, such as jellies, have elaborate gastrovascular cavities
Flatworms have a gastrovascular cavity and a large surface area to volume
ratio
Open and Closed Circulatory Systems
Complex animals have either open or closed circulatory systems
Both systems have three basic components:
A circulatory fluid (blood or hemolymph)
A set of tubes (blood vessels)
A muscular pump (the heart)
Open Circulatory Systems
In insects and other arthropods, and most molluscs, blood
bathes the organs directly in an open circulatory system
In an open circulatory system, the general body fluid is called
hemolymph
Closed Circulatory Systems
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In a closed circulatory system, blood is confined to vessels and is
distinct from the interstitial fluid
Closed systems are more efficient at transporting circulatory fluids
to tissues and cells
Organization of Vertebrate Circulatory Systems
Humans and other vertebrates have a closed circulatory system, often
called the cardiovascular system
The three main types of blood vessels are arteries, veins and
capillaries
Arteries branch into arterioles and carry blood to capillaries
(arterial system)
Networks of capillaries called capillary beds are the sites of
chemical exchange between the blood and interstitial fluid
Venuoles converge into veins and return blood from capillaries to
the heart (venous system)
General Goals of Circulation
Gas Exchange
Deliver respiratory gases to the tissues in the body for cellular
respiration
Remove the gases that are a byproduct of cellular respiration to the
respiratory surfaces for disposal
Energy Balance
Deliver the byproducts of digestion and absorption to the liver and
other body tissues for processing
Remove the waste products of nutrient breakdown during cellular
respiration to the kidney for excretion
Osmoregulation
Carry water, other ions, and molecular signal molecules throughout
the body
Different ways for the vertebrate heart to meet the goals of circulation in different
vertebrate classes
Gas Exchange (two plans)
Single circuit: pumps blood to the respiratory surfaces so it can
dump CO2 and pick up O2 for delivery directly to tissues; low
pressure throughout
Two circuits
:
First pumps blood at low pressure
to the lungs for
exchange of gases
Second pumps blood at high pressure
to the body for
delivery of gases to tissues (and takes CO2 to the heart and
then lungs)
Energy Balance (two results)
Single circuit: low pressure means slow delivery
. This is found in
animals with relatively low metabolic rates
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Document Summary

All organisms must exchange materials with the environment. The exchanges of gases used in cellular respiration (o 2 and co 2 ) is one of the most important processes in an animal"s physiology. Most gases move easily through membranes by diffusion. Although diffusion requires no energy, it is a random process and is relatively. Therefore, diffusion is fine for single celled and very small animals. Larger animals must build distributive systems for the transport of gas and other slow materials. Distributive systems take advantage of convection ( bulk flow ) through circulating blood to carry materials from one place to another in the body quickly and efficiently. Circulatory systems link exchange surfaces with cells throughout the body surrounding medium. In small and/or thin animals, cells can exchange materials directly with the. In most animals, transport systems connect the organs of exchange with the body. Most complex animals have internal transport systems that circulate fluid.

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